This is RAxML version 8.2.12 released by Alexandros Stamatakis on May 2018. With greatly appreciated code contributions by: Andre Aberer (HITS) Simon Berger (HITS) Alexey Kozlov (HITS) Kassian Kobert (HITS) David Dao (KIT and HITS) Sarah Lutteropp (KIT and HITS) Nick Pattengale (Sandia) Wayne Pfeiffer (SDSC) Akifumi S. Tanabe (NRIFS) Charlie Taylor (UF) Alignment has 5341 distinct alignment patterns Proportion of gaps and completely undetermined characters in this alignment: 4.05% RAxML Model Optimization up to an accuracy of 0.100000 log likelihood units Using 1 distinct models/data partitions with joint branch length optimization All free model parameters will be estimated by RAxML GAMMA model of rate heterogeneity, ML estimate of alpha-parameter GAMMA Model parameters will be estimated up to an accuracy of 0.1000000000 Log Likelihood units Partition: 0 Alignment Patterns: 5341 Name: No Name Provided DataType: AA Substitution Matrix: WAG Using fixed base frequencies Overall Time for Tree Evaluation 82.660923 Final GAMMA likelihood: -795835.602795 Number of free parameters for AIC-TEST(BR-LEN): 796 Number of free parameters for AIC-TEST(NO-BR-LEN): 1 alpha[0]: 0.895619 Tree-Length: 35.901365 rate A <-> R: 0.705217 rate A <-> N: 0.651871 rate A <-> D: 0.944853 rate A <-> C: 1.313132 rate A <-> Q: 1.161697 rate A <-> E: 2.023768 rate A <-> G: 1.811361 rate A <-> H: 0.405245 rate A <-> I: 0.247190 rate A <-> L: 0.508758 rate A <-> K: 1.158714 rate A <-> M: 1.142388 rate A <-> F: 0.269129 rate A <-> P: 1.839272 rate A <-> S: 4.309757 rate A <-> T: 2.711966 rate A <-> W: 0.144647 rate A <-> Y: 0.307794 rate A <-> V: 2.564804 rate R <-> N: 0.812328 rate R <-> D: 0.188337 rate R <-> C: 0.675324 rate R <-> Q: 3.881068 rate R <-> E: 0.561488 rate R <-> G: 0.747529 rate R <-> H: 2.732474 rate R <-> I: 0.239064 rate R <-> L: 0.636302 rate R <-> K: 6.842111 rate R <-> M: 0.873463 rate R <-> F: 0.131322 rate R <-> P: 0.868767 rate R <-> S: 1.565200 rate R <-> T: 0.708850 rate R <-> W: 1.488141 rate R <-> Y: 0.487813 rate R <-> V: 0.322004 rate N <-> D: 6.941838 rate N <-> C: 0.339146 rate N <-> Q: 1.973636 rate N <-> E: 1.211049 rate N <-> G: 1.439096 rate N <-> H: 5.058353 rate N <-> I: 0.708624 rate N <-> L: 0.168166 rate N <-> K: 3.851035 rate N <-> M: 0.253437 rate N <-> F: 0.122949 rate N <-> P: 0.249423 rate N <-> S: 5.081291 rate N <-> T: 2.595553 rate N <-> W: 0.091950 rate N <-> Y: 1.388516 rate N <-> V: 0.250912 rate D <-> C: 0.038734 rate D <-> Q: 0.788594 rate D <-> E: 7.894033 rate D <-> G: 1.106701 rate D <-> H: 1.189925 rate D <-> I: 0.050423 rate D <-> L: 0.108428 rate D <-> K: 0.613523 rate D <-> M: 0.132656 rate D <-> F: 0.059748 rate D <-> P: 0.542089 rate D <-> S: 1.370309 rate D <-> T: 0.479289 rate D <-> W: 0.165915 rate D <-> Y: 0.416441 rate D <-> V: 0.194769 rate C <-> Q: 0.126345 rate C <-> E: 0.027300 rate C <-> G: 0.392101 rate C <-> H: 0.318326 rate C <-> I: 0.217528 rate C <-> L: 0.491334 rate C <-> K: 0.094657 rate C <-> M: 0.499255 rate C <-> F: 0.508892 rate C <-> P: 0.139880 rate C <-> S: 1.799778 rate C <-> T: 0.655881 rate C <-> W: 0.916817 rate C <-> Y: 0.695323 rate C <-> V: 1.281296 rate Q <-> E: 6.993045 rate Q <-> G: 0.421991 rate Q <-> H: 5.490277 rate Q <-> I: 0.145650 rate Q <-> L: 1.111694 rate Q <-> K: 4.979863 rate Q <-> M: 1.975707 rate Q <-> F: 0.127755 rate Q <-> P: 1.193372 rate Q <-> S: 1.315472 rate Q <-> T: 1.096912 rate Q <-> W: 0.275833 rate Q <-> Y: 0.291141 rate Q <-> V: 0.385206 rate E <-> G: 0.725860 rate E <-> H: 0.728811 rate E <-> I: 0.162882 rate E <-> L: 0.197234 rate E <-> K: 3.304348 rate E <-> M: 0.402905 rate E <-> F: 0.103735 rate E <-> P: 0.872432 rate E <-> S: 0.901307 rate E <-> T: 1.051954 rate E <-> W: 0.200167 rate E <-> Y: 0.250985 rate E <-> V: 0.752728 rate G <-> H: 0.318886 rate G <-> I: 0.038932 rate G <-> L: 0.078380 rate G <-> K: 0.477616 rate G <-> M: 0.222597 rate G <-> F: 0.063840 rate G <-> P: 0.311419 rate G <-> S: 1.715597 rate G <-> T: 0.288741 rate G <-> W: 0.430853 rate G <-> Y: 0.132464 rate G <-> V: 0.239406 rate H <-> I: 0.176684 rate H <-> L: 0.638592 rate H <-> K: 1.138471 rate H <-> M: 0.516718 rate H <-> F: 0.868616 rate H <-> P: 0.890131 rate H <-> S: 0.946350 rate H <-> T: 0.605151 rate H <-> W: 0.335710 rate H <-> Y: 4.952425 rate H <-> V: 0.151328 rate I <-> L: 4.054275 rate I <-> K: 0.414039 rate I <-> M: 5.443417 rate I <-> F: 1.354596 rate I <-> P: 0.127765 rate I <-> S: 0.408423 rate I <-> T: 1.864345 rate I <-> W: 0.271672 rate I <-> Y: 0.537212 rate I <-> V: 10.000000 rate L <-> K: 0.329299 rate L <-> M: 6.206155 rate L <-> F: 2.704371 rate L <-> P: 0.531681 rate L <-> S: 0.440769 rate L <-> T: 0.417606 rate L <-> W: 0.850637 rate L <-> Y: 0.509657 rate L <-> V: 2.301842 rate K <-> M: 1.194528 rate K <-> F: 0.113582 rate K <-> P: 0.712025 rate K <-> S: 1.236534 rate K <-> T: 1.773337 rate K <-> W: 0.175808 rate K <-> Y: 0.170386 rate K <-> V: 0.390516 rate M <-> F: 1.522292 rate M <-> P: 0.219054 rate M <-> S: 0.631487 rate M <-> T: 1.938450 rate M <-> W: 0.659361 rate M <-> Y: 0.547782 rate M <-> V: 2.631851 rate F <-> P: 0.206416 rate F <-> S: 0.698005 rate F <-> T: 0.219788 rate F <-> W: 1.955736 rate F <-> Y: 8.252183 rate F <-> V: 0.830926 rate P <-> S: 2.062675 rate P <-> T: 1.016946 rate P <-> W: 0.178238 rate P <-> Y: 0.276228 rate P <-> V: 0.402602 rate S <-> T: 5.597561 rate S <-> W: 0.669635 rate S <-> Y: 1.006218 rate S <-> V: 0.297571 rate T <-> W: 0.141746 rate T <-> Y: 0.372250 rate T <-> V: 1.774935 rate W <-> Y: 3.177720 rate W <-> V: 0.467146 rate Y <-> V: 0.402401 freq pi(A): 0.086628 freq pi(R): 0.043972 freq pi(N): 0.039089 freq pi(D): 0.057045 freq pi(C): 0.019308 freq pi(Q): 0.036728 freq pi(E): 0.058059 freq pi(G): 0.083252 freq pi(H): 0.024431 freq pi(I): 0.048466 freq pi(L): 0.086209 freq pi(K): 0.062029 freq pi(M): 0.019503 freq pi(F): 0.038432 freq pi(P): 0.045763 freq pi(S): 0.069518 freq pi(T): 0.061013 freq pi(W): 0.014386 freq pi(Y): 0.035274 freq pi(V): 0.070896